Postdoctoral Fellow, Albert Einstein College of Medicine, Bronx, NY, Molecular Biology (1967)
Senior Resident, Duke University Hospital, Durham, NC, Medicine (1965)
Clinical Associate, National Institute for Arthritis and Metabolic Diseases, National Institutes of Health, Bethesda, MD, Molecular Biology (1964)
Senior Assistant Surgeon, Reserve Corps of the Public Health Service, Molecular Biology (1964)
Resident, University of Michigan, Ann Arbor, MI, Medicine (1962)
Intern, Mount Sinai Hospital, New York, NY, Medicine (1961)
M.D., University of Pennsylvania, Philadelphia, PA, Medicine (1960)
Summer Student, Laboratory of Peter Medawar, University College London, UK, Transplantation Immunology (1959)
B.S., Rutgers University, New Brunswick, NJ, Biological Sciences (1956)